Tjio, Joe‐Hin

Maj A Hultén, University of Warwick, Coventry, UK

Published online: September 2006

DOI: 10.1002/9780470015902.a0006252

Joe-Hin Tjio (1919–2001) was a Java-born scientist who, in collaboration with Albert Levan, uncovered the correct number of human chromosomes.

Keywords: chromosome number; cytogenetics; Levan

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Further Reading
	Hultén MA (2002) Numbers, bands and recombination of human chromosomes: historical anecdotes from a Swedish student. Cytogenetics and Genome 96: 14–19.
	Levan A (1978) The background of the determination of the human chromosome number. American Journal of Obstetrics and Gynecology 130: 725–726.
	Tjio J-H and Levan A (1956) The chromosome number of man. Hereditas 42: 1–6.
	Tijo JH (1978) The chromosome number of man. American Journal of Obstetrics and Gynecology 130: 723–724.

	Figure 1. Joe-Hin Tjio (1919–2001), ca. 1960.
	Figure 2. Photograph taken at the ceremony of the Kennedy International Award in 1962. Tjio is standing to the left of Jerome Lejeune. (Photo courtesy of Tjio.)
	Figure 3. Examples of the microphotographs taken by Tjio. On each photograph he has written, in a combination of Chinese and Swedish, ‘to Maj Hultén with affection’ followed by the species, the time the photo was taken, and the time it was given to her. (a,b) Metaphase plates of human chromosomes; (c) Ehrlich ascites tumor metaphase; (d) Yoshida sarcoma (rat) quadripolar anaphase; (e) Luzula purpurea (2N=6) late prophase, diffuse centromere.